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Would you like to inspect the original subtitles? These are the user uploaded subtitles that are being translated: 0 00:00:00,000 --> 00:00:00,792 MICHAEL HEMANN: OK. 1 00:00:00,792 --> 00:00:03,840 So how do we figure out what the double recombination 2 00:00:03,840 --> 00:00:05,850 events are? 3 00:00:05,850 --> 00:00:08,430 Well, essentially, we just try out 4 00:00:08,430 --> 00:00:10,590 what things would look like if they 5 00:00:10,590 --> 00:00:13,930 were in all of the different possible orders. 6 00:00:13,930 --> 00:00:16,350 So there are three orders that could possibly 7 00:00:16,350 --> 00:00:19,350 exist amongst these genes. 8 00:00:19,350 --> 00:00:21,300 We're going to model these double crossovers. 9 00:00:21,300 --> 00:00:25,410 And we're going to see which one appears 10 00:00:25,410 --> 00:00:30,000 to be the double crossover based on the numbers that we see, 11 00:00:30,000 --> 00:00:31,320 right? 12 00:00:31,320 --> 00:00:41,620 So let's look at order number one, right? 13 00:00:41,620 --> 00:00:47,050 Say we have wingless minus M1A M2A. 14 00:00:47,050 --> 00:00:49,510 Now, that's the way I've drawn it before. 15 00:00:49,510 --> 00:00:51,820 But now we're saying that this is actually 16 00:00:51,820 --> 00:00:55,760 the order that they're in on the chromosome. 17 00:00:55,760 --> 00:00:57,400 So we have that. 18 00:00:57,400 --> 00:01:06,520 And we have, then, wingless plus M1B M2B. 19 00:01:06,520 --> 00:01:08,470 So we're going to have-- 20 00:01:08,470 --> 00:01:12,260 21 00:01:12,260 --> 00:01:15,810 what would happen if there's a double crossover there, right? 22 00:01:15,810 --> 00:01:19,170 23 00:01:19,170 --> 00:01:22,230 So that recombination product that you would get 24 00:01:22,230 --> 00:01:36,930 would be wingless minus M1B M2A and wingless plus M1A M2B. 25 00:01:36,930 --> 00:01:40,737 26 00:01:40,737 --> 00:01:41,570 How is this working? 27 00:01:41,570 --> 00:01:44,640 Well, I'm starting over here at wingless. 28 00:01:44,640 --> 00:01:49,020 I'm going down to M1B, and I'm going back up to M2A. 29 00:01:49,020 --> 00:01:51,810 So you're getting wingless minus M1B M2A. 30 00:01:51,810 --> 00:01:55,490 31 00:01:55,490 --> 00:02:07,090 The gametes are wingless minus M1B M2A and wingless plus M1A 32 00:02:07,090 --> 00:02:07,590 M2B. 33 00:02:07,590 --> 00:02:10,340 34 00:02:10,340 --> 00:02:10,840 OK. 35 00:02:10,840 --> 00:02:13,420 So that's one possible order. 36 00:02:13,420 --> 00:02:17,320 37 00:02:17,320 --> 00:02:29,400 The second possible order would be M1A wingless minus M2A 38 00:02:29,400 --> 00:02:34,810 and M1B wingless plus M2B, right? 39 00:02:34,810 --> 00:02:37,857 40 00:02:37,857 --> 00:02:39,940 Again, we think about what would happen if there's 41 00:02:39,940 --> 00:02:41,260 a double crossover here. 42 00:02:41,260 --> 00:02:47,070 43 00:02:47,070 --> 00:02:56,400 And the result would be M1A wingless plus M2A. 44 00:02:56,400 --> 00:02:59,282 45 00:02:59,282 --> 00:03:05,860 And you would get M1B wingless minus M2B. 46 00:03:05,860 --> 00:03:08,560 47 00:03:08,560 --> 00:03:10,570 And the gametes-- we can write out-- 48 00:03:10,570 --> 00:03:12,760 in this case, I'm not writing out the order. 49 00:03:12,760 --> 00:03:16,285 I'm just writing out based on the same sort of convention 50 00:03:16,285 --> 00:03:19,810 of what the gametes would look like from a genetic standpoint. 51 00:03:19,810 --> 00:03:26,340 So this gamete would be wingless plus M1A M2A 52 00:03:26,340 --> 00:03:32,385 and wingless minus M1B M2B. 53 00:03:32,385 --> 00:03:36,240 54 00:03:36,240 --> 00:03:42,450 Let's think about a third order here. 55 00:03:42,450 --> 00:03:45,030 And the third order would be the final order, 56 00:03:45,030 --> 00:03:57,575 which would be M1A M2A wingless minus and M1B M2B wingless 57 00:03:57,575 --> 00:03:58,075 plus. 58 00:03:58,075 --> 00:04:02,530 59 00:04:02,530 --> 00:04:05,670 I doublecross over here, right? 60 00:04:05,670 --> 00:04:27,620 And the results would be M1A M2B wingless minus and M1B M2A 61 00:04:27,620 --> 00:04:30,020 wingless plus, right? 62 00:04:30,020 --> 00:04:38,040 And we write those all with the same sort of convention here. 63 00:04:38,040 --> 00:04:41,370 We have wingless minus M1A M2B. 64 00:04:41,370 --> 00:04:48,170 And we have wingless plus M1B M2A. 65 00:04:48,170 --> 00:04:49,000 4324

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